This book is about the interrelationship between nature semiosis metarepresentation and selfconsciousness and the role played by metarepresentation in evolution. Representations must have emerged via selforganization from nonrepresentational systems found in physics chemistry and biology. Major steps have been the evolution of molecules macromolecules life and finally cultural and symbolic systems. Representations and signs are therefore parts of a huge possibly branching ladder of beings. Metarepresentations images representing images language about language and languageuse thoughts about thoughts constitute a fascinating theme within such diverse areas of research as philosophy literature theology anthropology and history neuroscience psychology and linguistics. The contributions to this book reflect this variety of different but often interrelated perspectives on metarepresentation. They also exemplify the difficulties of a truly interdisciplinary discourse and show how one may start such a discourse in the field of semiotics understood as a metadiscipline which brings together all scientific enterprises dealing with human mind and human culture.